第十三章

<= /div>

<= b>页眉

<= /div>

‹<= b>日期/<= b>时间›

<= /div>

̸= 9;<= b>页脚›

<= /div>

‹#›

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第三篇 基因信息௚= 0;传递

复= 1046;

翻= 5793;

蛋白质

（病毒）

RNA

（病毒）

= 逆转录

转录

RNA

翻译

蛋= 0333;质

DNA

复制

中心法则（The central dogma）:

基因：为生物活性产物编码= ;的DNA功能片段

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第十一章

DNA的生物合成

复制：遗传&= #20449;息从亲代DNA传递到子代 = DNA分子上。

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第一节    半保留复制

(Semiconservative Replication)

定义：子&#= 20195;DNA双链中的一条链来&#= 33258;
      母链，另一条链重= 6032;合成。

母链

子链

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(<= /span>深蓝: = 15N)<= /span>

(<= /span>粉红: 14N)<= /span>

DNA半&#= 20445;留复制的证据

培养&= #20110;普通培养液

继续培= 养于

普通&= #22521;养液

含15N-DNA的细菌

= 普通DNA=

重DNA

第一代=

中等密度DNA

第二代

<= b>

= 普通DNA=

中= 1561;密度DNA

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重<= /span>DNA

普通DNA

中等密度= DNA

全保留= 式

混合式=

第一代

第二代

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第一代

第二代=

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------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/slide0049.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" 第十三章

半保= 留复制的意义：
遗传= 稳定性的分子机制。

但是：遗传稳定性是相对= = 的，生物஺= 8;存在普遍的变异= 9616;象

基因表达：包括转录和翻译

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第二节 DNA复制的酶&#= 23398;

1. 底物：dATP，dGTP，dCTP，= dTTP， <= /span>总称为dNTP

2. DNA聚合酶，DNA-po= l

3. 模板（templa= te）：解开成单链的DNA母链

4. 引物（primer= ）：RNA引物

5. 其他酶和蛋白质因子= ;

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一、复制的化= 学反应

5`=

3`=

3`=

5`=

磷酸二酯键的生成

3`=

5`=

N1OH=

3`

5`

+d= N2TP=

N1N2-OH

3`

5`

+PPi

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DNA pol

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=

=

二、= DNA聚合酶<= /b>（DNA polymerase, DNA-pol）

     即依赖于DNA的DNA聚合Ŵ= 38;（DDDP）

=

=

y真核生物有多种： = ;

        = ;      DNA-pol a、b= 289;g、d、e…

y原核生= ;物有3种： <= /span>

        = ;      DNA-polⅠ、 Ⅱ、 Ⅲ

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   表<= /b>13-1    大&= #32928;杆菌中DNA聚&= #21512;酶某些性质

        = ;            &n= bsp;            = ;            &n= bsp;            = ;    DNA-pol

    性质        = ;            &n= bsp;     Ⅰ        = ;      Ⅱ        = ;     Ⅲ
酶分子/细&= #32990;        = ;             400        = ;    40        = ;      20

活性（nt/min<= /span>）<= /b>        = ;      1000        = ; 50     &n= bsp;    100000

5 ' →3 'ň= 58;合功能        = ;        有        = ;   有        = ;     有

5 ' →3 '外切酶活&= #24615;        = ;   有        = ;    有        = ;     <= b>无

3 ' →5 '外切酶活&= #24615;        = ;   有        = ;    有        = ;     有

主要功能<= /b>        = ;            校读 ,修&= #22797;   不&= #28165;         <= b>复制

        = ;            &n= bsp;            = ;    填补缺口

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a

a

t

t

e

e

q

q

b

b
=
b

c

g

d’ <= /div>
j

d= 3;

b

The model of DNA-pol III

a e<= span style=3D'font-size:150%;color:#FF3300'> q<= span style=3D'font-size:117%'> form the catalytic core

t Links two cores

Leading strand = synthe= sis

Lagging strand = synthe= sis

b act as a clamp

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小片段

=         = ;      大片段

（Klenow fragment）

5 ‘ →3 ’<= b>聚合功能，

3 ' →5 '外切酶活&= #24615;

5 ' →3 '外切酶活&= #24615;

DNA pol I

12/62

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charset="us-ascii" 第十三章

真核细= 2990;中DNA聚合酶

种类： DNA-pol α、β、γ、δ、ε…

DNA-pol α<= /b>：合成随从链=

DNA pol δ<= /b>：合成领头链=

DNA-pol ε<= /b>：校读，修复和= 2635;补缺口

DNA-pol γ<= /b>: ߠ= 5;存在于线粒体中

DNA-pol β<= /span>：功能不清

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------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/slide0085.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" 第十三章

     质粒与线粒体DNA是细胞核染色体外<= span style=3D'font-family:SimSun;mso-fareast-font-family:SimSun;font-size:133%'= >的遗传物质。能进行= ;自我复制。

质粒是基因= 工程的常用载体

线粒体DNA(mtDNA)全长16kb，有37= 个基= 因，编码与ATP合成有关的酶和蛋白= ;质。

特点：

1. 母系遗传

2. DNA= 损伤后不易修复

3. 遗传密码与通用遗传= ;密码有差别

14/62

UGA（终止密码子）：Trp

AGA/AGG（Arg）：终止密码子

AUA（Ile）：Met（起始密码子）=

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DNA-pol = 0340; 5´= ®3= ´聚= ;合作用

3’<= /span>

5’<= /span>

5’<= /span>

3’<= /span>

3’<= /span>

5’<= /span>

DNA-pol

15/62

5’<= /span>

3’<= /span>

OH

P

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3´®5´外= ;切 <= /div>

5´® 3´= 外切

5’

3’

内切酶

（限制性内切酶= ;）

5̵= 7;

3̵= 7;

3´®5´外= ;切 <= /div>

5´® 3´= 外切

外切酶与内切酶= 作用图解

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DNA-pol <= /span>I <= /span>5´® = 3´外切酶活性的功= 能：

切除引物，切除突变= ;片段

DNA-pol <= /span>I <= /span>3´®5= ´外切酶活性的功= 能：

校读（proofread）功能

5’<= /span>

3’<= /span>

A

G

5’<= /span>

3’<= /span>

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DNA复制的保真性(fidelity)

1.严格遵守碱基配= 对规律

2. DNA-pol在复制过&#= 31243;中对碱基的选择性<= /span>

3. 复制出错时&= #26377;即时的校读功能<= /div>

A

T

G

C

DNA pol III在催化磷酸二酯= ;键形成之前完成对碱&#= 22522;<= span style=3D'font-family:SimSun;mso-fareast-font-family:SimSun;font-size:117%'= >的选择；对反式嘌呤核苷酸= 的亲和力较顺式的大= ;。

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三、复制中解链= 和DNA分子的拓扑学变= 化

解链酶类：

1. 解螺旋酶(helicase)

2.拓扑异构酶（topoisomerase I、II）=

=

3. 单链DNA结合蛋白

(sin= gle stranded DNA-binding protein,SSB)

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1.Š= 99;螺旋酶 (helicase)

解螺旋酶

作用：断裂= 114;补碱基间的氢键，使= ;DNA成单链。

dnaA、B、C<= /span>…= ;

DnaA、B、C<= /span>…= ;

ATP

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2. 拓扑异构酶（topoisomerase,Topo）

拓扑(topol= ogy)与拓扑异构

DNA正超螺旋与负超螺旋= ;

负超螺旋

正超螺旋

DNA双螺旋

拓扑异构&#= 37238;

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切割<= span lang=3DEN-US style=3D'font-family:"Times New Roman";mso-ascii-font-family:= "Times New Roman"; mso-fareast-font-family:KaiTi_GB2312;font-size:133%;mso-fareast-language:Z= H-CN'>DNA双໦= 2;，此时<= span lang=3DZH-CN style=3D'font-family:KaiTi_GB2312;mso-fareast-font-family:Kai= Ti_GB2312; font-size:133%;color:#3333FF;mso-fareast-language:ZH-CN'>不༙= 6;ATP；尔后= 由A= TP供能，使<= span lang=3DEN-US style=3D'font-family:"Times New Roman";mso-ascii-font-family:= "Times New Roman"; mso-fareast-font-family:KaiTi_GB2312;font-size:133%;mso-fareast-language:Z= H-CN'>DNA分子成= 负超&#= 34746;旋= 再连接= 切口。 =

不需ATP，切割双链DNA中的= 968;链ʌ= 92;使DNA松弛后, 连接切口。

Topo= Ⅰ:

Topo= Ⅱ:

      临床上使用的某些抗= ;肿瘤药（如喜树= = 碱，鬼臼乙叉甙等）= ;是通过抑制Topo<= /b>酶活性= 而杀死肿瘤细胞的。= ;

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Topo酶：切割DNA链，使其松弛后再连= ;接。

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=

3. 单链DNA结合蛋白（single stranded binding protein，SSBA= 289;=

作用：防止单链DNA重新形= 成双链，= ༅= 0;止单链DNA被核酸酶水෉= 9;。

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Content-Transfer-Encoding: base64 Content-Type: image/gif R0lGODlhKAAEAHcAMSH+GlNvZnR3YXJlOiBNaWNyb3NvZnQgT2ZmaWNlACH5BAEAAAAALAMAAAAj AAQAgAAAAP8zAAIKjI+py+0Po5wUFQA7 ------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/slide0015.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" 第十三章

=

=

= 2235;、引物酶(Primase)和引发体= ;

= 5´

5´

催化RNA引物<= span style=3D'font-family:KaiTi_GB2312;mso-fareast-font-family:KaiTi_GB2312; font-size:133%'>合成的酶叫引物酶，它是一种特殊的RNA聚合酶。

DNA合成需在RNA引物的基础= 上进行。

<= ![if !ppt]>
RNA引&#= 29289;

5´

3<= /span>´

5´

3<= /span>´

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DnaA

引发体（primosome）：包括解螺旋酶、DnaC、引物酶及DNA复制&= #30340;起始区域。

5´

3<= /span>´

3<= /span>´

5´

引物酶

DnaC

引发体

解螺旋酶

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file:///C:/85899A51/11.files/slide0067_image116.gif Content-Transfer-Encoding: base64 Content-Type: image/gif R0lGODlhygBDAHcAMSH+GlNvZnR3YXJlOiBNaWNyb3NvZnQgT2ZmaWNlACH5BAEAAAAALAIADgBY ADEAgAAAAAAAAAKUhI+py+0PWQix2ovbzLx7tX3iWE0UiaaIqbZp6MYeLNcXbecPq/cO7gsagEIf sajjIYPKpfHkfEaZ0GmuaZVhs7Ej91X9urzi0baMIqNn4TVJ7c6c4505nQO/l9p6PL+PYQcYITgI kWco8Zd4uMj4g/i44ii5EFkJcFmpKVmIOfm5F6pxwGlo4vmJOrqzyqqISvlZAAA7 ------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/slide0068.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" 第十三章

五、 DNA连接酶(DNA ligase)

=

=

=

功能： =

=

=

=

      催化DNA双链的<= /span>3’羟= ;基和相邻的5’磷= ;酸= 基团形成磷酸二酯键= ，从而把两段= 相邻= 的DNA链连成完ă= 72;的链。

=

=

=

=

=

=

DNA连接Ŵ= 38;

=

=

=

ATP

ATP

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        = ;

<= /b>

<= /b>

<= /b>

参与= DNA复制的= 0027;要成员

主要成= 1592;

主要作= 9992;

DnaA        = ;       <= b>识别复制起始位点

解螺旋酶<= b>    解开DNA双&= 142;

SSB        = ;            &n= bsp; <= b>维持已解开单链DNA的稳定

引物酶        = ;     合= 104;RNA引物

TOPO        = ;   <= b>使打结、缠绕、正超&= #34746;旋的DNA松驰=

DNA-pol Ⅲ     DNA<= span style=3D'font-family:SimSun;mso-fareast-font-family:SimSun;mso-hansi-font-= family: "Courier New";font-size:133%'>复制

DNA-polⅠ=      水解引= 9289;、填补空隙、修复߯= 6;用

DNA连接酶    催化双链DNA中单链缺口的连接<= /b>

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   <= b>

第三节DNA生物合成过= ;程

1.复制的起&#= 22987;

2.链的ॿ= 0;长

3.复制௚= 0;终止

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S

G1

M

G2

蛋白质合成期

DNA合成期

哺乳动= 物的细胞周期

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复制= ;的起始

=

=

(一)DNA解成单&= #38142;

由特定蛋白质识别= 2797;制起始位点（ori），= 在解螺旋Ŵ= 38;、TOPO酶及单链DNA结合蛋&= #30333;的共= 同作用下ʌ= 92;DNA解链，解旋，形成复制&#= 21449;。

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q <= /div>

倒Y

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E.coli= 复制起始点oriC跨度为245bp<= /b>，有= 3<= /span>组串联重复序列和2<= /span>对反向重复序列。

GATTNTTTATTT…GATCTNTTNTATT…GATCTCTTATTAG

串联重复序列

反向重复序列

= …TTTGGATAA…..<= span lang=3DEN-US style=3D'color:red;mso-fareast-language:ZH-CN'>TTATCCA= CA
        = ;            &n= bsp;            = ; AATAGGT--T

= TGTGGATTA……TTATACACA…

        = ;            &n= bsp;            A--TA<= /span>--GTGT

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回文结构

AGCT&#= 13;

TCGA

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(二)引发体的ஶ= 3;成

解螺旋酶解开双链= 1518;引物酶进入形成引发= 体。 <= /span>

5´

3<= /span>´

3<= /span>´

5´

引物酶

DnaC

引发体

解螺旋酶

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text/html; charset="us-ascii" 第十三章

（三） RNA引物į= 40;合成<= b>

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<= ![if !ppt]>
= 5´

5´

RNA引&#= 29289;

5´

3<= /span>´

5´

3<= /span>´

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小节

参与复= 制起始的成员

名称

功能

DnaA蛋白        = ;            &n= bsp;          辨认起始点

解螺旋酶（DnaB，Rep）解开DNA双= ;链

DnaC        = ;            &n= bsp;            = ;    协助解螺旋酶

引&#= 29289;酶（DnaG）        = ;       催化RNA引物生成

SSB        = ;            &n= bsp;            = ;       稳定解开的单= 链

拓&#= 25169;异构酶        = ;            &n= bsp; 理顺DNA链

OriC        = ;            &n= bsp;            = ;     E.coli复制起始点

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     引物合成后A= 292;由DNA polⅢ（在真核= ;细胞<= span style=3D'font-family:KaiTi_GB2312;mso-fareast-font-family:KaiTi_GB2312; font-size:133%;color:blue'>为DNA聚合酶d和= ;a)催化，按= ;碱基配对原则，= <= span style=3D'font-family:KaiTi_GB2312;mso-fareast-font-family:KaiTi_GB2312; font-size:133%;color:blue'>将dNTP逐一添加= ;到引物3’ 末端，形成<= span style=3D'font-family:KaiTi_GB2312;mso-ascii-font-family:KaiTi_GB2312;mso-f= areast-font-family: KaiTi_GB2312;font-size:133%;color:#FF3300'>磷酸<= /span><= span style=3D'font-family:KaiTi_GB2312;mso-ascii-font-family:KaiTi_GB2312;mso-f= areast-font-family: KaiTi_GB2312;font-size:133%;color:#FF3300'>二酯键A= 292;使新合成的&= 142;不断延长。

复= 046;的延长

3<= /span>’= ;——AAGACC= TATT—= ;—5<= /span>’= ;

5̵= 7;——TTCTGGATAA—= ;—3<= /span>’= ;

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= <= ![endif]>

dATP

+<= /span>

DNA pol

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领头链= 5288;leading strand）

随从链= 5288;lagging = strand）

<= span style=3D'font-family:KaiTi_GB2312;mso-fareast-font-family:KaiTi_GB2312; font-size:133%;color:#FF3300'>冈崎片段= (Okazaki fragment)

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1.半不连续性：DNA复制过程中，一Ĉ= 65;链是连续复制，另一&= #26465;链是不连续复制的= 616;象。

有关复制延&#= 38271;的几个重要概念

4. = 冈崎片段（<= /b>Okazaki fragmentA= 289;：

      不连续复制&= #30340;片段

3. 随从链: 链的延长方&= #21521;(5’®3’)与解链= 041;向= 相反，为不&= #36830;续复制。

2.领头链:&= 142;的延长方向(5’® 3’)与解链= 041;向<= span style=3D'font-family:"Times New Roman";mso-ascii-font-family:"Times New Ro= man"; mso-fareast-font-family:KaiTi_GB2312;font-size:133%;color:#3333FF'> 相同, 为连续= 797;制。

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5. 双向复制=

ori

ori

6. 复制子(replicon)

ori

ori

真核生物两个相= 邻复制起始&= #28857;之= 间的DNA片段。=

以起始点为&= #20013;= 心，向两个&= #26041;= 向进行复制&= #12290;

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滚环复制

= 是某些病毒，质粒、= ;线粒体= DNA的特殊复制形式。

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复制的终止

•原&#= 26680;生物复制终止

•真&#= 26680;生物复制终止

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------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/slide0076.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" 第十三章

1、原核生物DNA为环状，双向复= 制的汇<= span style=3D'font-family:SimSun;mso-fareast-font-family:SimSun;font-size:133%; color:#3333FF'>合点就是复制终= 止点。

原核生物复= 1046;终止

2. 领头链上的<= /b>RNA引物被RNA酶水解后，由= DNA pol I 催化，由新&= #21512;成链提供-OH补齐&#= 12290;

终点

起点

RNA= = 酶

DNA pol= I

连= 5509;酶

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3、&#= 20872;崎片段引物的消除Ø= 44;连接。

RNA引物

RNA酶

DNA pol I

DNA连接酶

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真核生物的端粒(telomere)和= = 端粒酶(telomerase= )

真&#= 26680;生物DNA末端结构是端<= span style=3D'font-family:SimSun;mso-fareast-font-family:SimSun;font-size:133%; color:#3333FF'>粒，在端粒酶的= 作用下形= 成。

（TTAGGG= ）n<= /span>

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端粒酶：由RNA与蛋白质组成，= 具逆转录酶活 性，以自身RNA为模板合成DNA。

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DwwKLbmdqzdMKpdMVNHYQio+1Kr1is1qt9yu9wsOi8fk8hZ6bJjX7Lb7DY/Lv2he847P69Vo6f4P GPjz8eIheIiYKOOn2Oj4CBkpOUlZaXmJmam5ydnp+QkaKjpKWmp6iooRsMq6mvo6ugoVAFvbGVAn QGvLe4mbu9srHJnLEjyMrPhbFzDn/AwdLT0NlrA8S52tvc3d3aUga3ScTK7cylqejlgAADs= ------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/slide0058.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" 第十三章

U

U

A

A

3’= ;

5’= ;

A

C

<= span style=3D'mso-spacerun:yes'> CAACC

C

GGGTTGGGT

3’= ;

5’= ;

3’ = ;

C        = ;          CAACC

GGGTTGGGT= TGG

U

U

A

A

3’= ;

5’= ;

3’= ;

5’= ;

3’ = ;

A

C

C        = ;            &n= bsp;   ACCUU

GGGTTGGGT= TGG

A

A

3’= ;

5’= ;

3’= ;

5’= ;

3’ = ;

A

C

C

A

Binding

Elongation

Translocation

Telomerase=

45/62

爬行模型：Inchwo= rm（尺蠖）model<= /b>

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C        = ;            &n= bsp;   ACCUU

GGGTTGGGT= TGGAA

A

A

3’= ;

5’= ;

3’= ;

5’= ;

3’ = ;

A

C

C

A

Elongation

C        = ;            &n= bsp;   ACCUU

GGGTTGGGT= TGGAA

3’= ;

5’= ;

3’ = ;

5’ = ;

Primer

synthesis

CCCAACCCAACCUU

GGGTTGGGT= TGGAA

3’= ;

5’= ;

3’ = ;

5’ = ;

Primase

DNA polymerase

CCCAACC= CA

GGGTTGGGT= TGGAA

3’= ;

5’= ;

3’ = ;

5’ = ;

DNA

replication

Primer

removal

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= 第四节 DNA损伤（突变）与修= 复

   DNA分子一级结构的= 改变称为DNA损= 伤(DNA damage)或突变（mutati= on）。

大多数突变属此类，= ;原因不明

由理化及生物因素诱= ;发

DNA的损伤= （突变）的概念

自发突变：

诱发突变：

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遗传重组不属突变的范畴。

------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/slide0037_image222.gif Content-Transfer-Encoding: base64 Content-Type: image/gif R0lGODlhQgEoAHcAMSH+GlNvZnR3YXJlOiBNaWNyb3NvZnQgT2ZmaWNlACH5BAEAAAAALAAAAABA AScAgQAAAJkz///t4QECAwLdjI+py+0Po5y02ouz3jyLD4biSJbmiabqyrbuC8fyTNevYef6zvf+ DwzScMKi8YhMKpcmIvMJjUqnTCf1is1qtyMr9wsOi3nesfmMToPK6rb7HWXD5/Q6OWDP6/cxOf8P uOcXSFjYNmiYqPiFuOj4GIcHOUkp1ViJmalzqdnp6cL5KTpKEkp6OmqKuqqpyvo66Qo7qyhLexto i7urp8v7O+cLPKwmTHw8Zoy8zKXM/HzlDD0NJU19nWSNvS2kzf3d4w0+biNOft4nib5erc7+nt0h P09fb3+Pn6/fUQAAOw== ------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/slide0044.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" 第十三章

一、基因突&#= 21464;的意义

（一）突变是进化、= ;分化的分子基础

（二）只有基因型改= ;变的突变

（三）致死性的突变= ;

（四）突变是某些疾= ;病的发病基础

同义突变

某些非编码区的突变= ;

DNA多态性：个体间DNA序列的差异

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如遗传性疾病，高血= ;压，肿瘤，糖尿病等&#= 31561;。

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O

N

H

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O

C

H

3

R

P

O

N

H

N

O

C

H

3

R

O

N

H

N

O

C

H

3

R

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UV

二、引发突变的= 因素

物理因素

紫外线(产生胸腺嘧啶二= 2858;体，      ）

TT

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环丁基环

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化&#= 23398;因素

ୋ= 9;化剂：(<= /span>如氮芥类)<= /span>，使鸟嘌呤的 N烷基化后= = 脱落，成为无鸟嘌呤= ;的位点

亚Ĵ= 13;酸盐：使碱基氧化脱氨基= ;

C<= /span>→U

A<= /span>→I

G<= /span>→X

碱基类似物

如5-= 溴尿嘧啶(<= /span>5-BU)与T类似，但5-BU可与G配对

羟胺类：C被羟胺化后与A配对

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—= ;   A

—= ;5-BU

—= ;   G

—= ;5-BU

------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/slide0039_image231.gif Content-Transfer-Encoding: base64 Content-Type: image/gif R0lGODlhMAAKAHcAMSH+GlNvZnR3YXJlOiBNaWNyb3NvZnQgT2ZmaWNlACH5BAEAAAAALAEAAwAs AAUAgAAAAAAAAAIWhI+pyxfRopzvgYqz3rzbCYJVSFJIAQA7 ------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/slide0040.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii" 第十三章

C

O

HN

C

CH

CH

= =

N

= =

O

U

C

O

N

C

CH

CH

=

N

NH2

C

A

N

NH2 = ;

N

N

N

N

O

HN

N

N

I

N

OH

N

N

N

H

H2N=

G

N

OH

N

N

N

H

HO

X

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三、突变的类型

1. 点突变（point mutation）

也称错配（mismatch）

DNA链上碱基的置换

CTC

Glu

CAC

Val

HbA b肽链

HbA b基因

HbS b基因

HbS b肽链

A

T

G

C

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2. 缺失、插入&= #21644;框移突变（frame shift）

5′…. UCAC= GACAUAUG….3′

5′….UCAG= CGACAUAUG….3′

丝

精

组

蛋

丝

丙

苏

酪

5′…. UCAGACAUAUG….3′

丝

天

异亮

插入

缺失

正常

框移突变是最严重的突= 变！

人之初，性本善，= 性相近，习相远…= ;

人初性，本善性，相= ;近习，相远…= ;

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3. 重排(rear= rangement)与重组(reco= mbination)

DNA分子发生较大片段的= ;交换

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四、DNA损伤的修复

光复活酶

（可见光）

T-T

1. 光修复

TT=

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= 2.切除修复：由3<= /span>种酶共同参= 与完成。

UvrC切除<= /div>
DNA pol I<= /span>

DNA= 连接酶

5'

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UvrA，UvrB识别并结合

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XP=

XP=

DNA po= l e

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3.重组修复

RecA

DNA pol

切除修复

57/62

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4. SOS修复：DNA分子多处受= 到较大范围的损<= span style=3D'position:absolute;top:57.14%;left:0%;width:100.0%;height:44.64%'>= <= b> 伤，细= 2990;所作出的一系列反ॵ= 2;。

细胞&= #33021;继续生存，但引起DNA较长期的= 广泛的突变。=

=
后果：

SOS修复系统：Uvr类，Rec类，DNA p= ol，LexA等

组成调节&#= 23376;（regulon）网络系统。

特点：抑制&= #32454;胞分裂，允许大量= 555;速的切除修<= span style=3D'font-size:133%;color:#3333FF'> 复和重&#= 32452;修复， 允许损= 伤的DNA作为模= = 板进行复制，对碱基选择性低= 等。

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第五节   <= /span>$= 870;转录现象和逆转录酶= ;

逆转录酶（reverse transcriptase）= 又= 称为反= ;转录酶，为依= ;赖RNA的= ;DNA聚= ;合酶= (RNA-dependent DNA polymerase, RDDP)

以RNA为模板= ;，dNTP为&#= 21407;料，按碱基配对规ô= 59;= 合&#= 25104;DNA的&#= 36807;程，由逆转录酶催&#= 21270;。<= b>

逆转录现ฉ= 7;

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逆转录酶&#= 26159;多功能酶，有三种&#= 37238;活性：

1. 逆&#= 36716;录活性：即以RNA为模= ;板合成DNA

2. RNase活性：水解RNA:DNA中的RNA

3. DNA pol活性：以DNA为模板合成DNA<= /span>

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RNA模板

逆转录酶&#= 30340;逆转录活性

逆转录酶&#= 30340;RNase活性

RNA:DN= A杂化双链

单链DNA

逆转录酶&#= 30340;DNA pol活性

整合(integration)= 入细胞基因组

双链DNA

     病毒基因组通过基因= 7325;组插入到宿主细胞基= 因组中，称为整合。

tRNA引物

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逆转录及逆转录= 酶的生物医学意义=

1. 丰富和发展&= #20102;中心法则

2. 丰富和发展&= #20102;致癌理论

3. 在基因工程&= #20013;的应用

RNA也可作为遗传信息的= ;载体

RNA在进化中可能比DNA更原始

癌基因与病&= #27602;致癌理论

逆转录酶广泛应用于= ;基因工程中

逆转录病毒是基因治疗的常用载体

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<= /div>
<= /div>
<= /div>

<= b>页眉

<= /div>
<= /div>
<= /div>

‹<= b>日期/<= b>时间›

单击此处编辑母版ă= 91;本样式

第二级

第三级

第四级

第五级

<= /div>
<= /div>
<= /div>

̸= 9;<= b>页脚›

<= /div>
<= /div>
<= /div>

‹#›

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g_bgSound=3D"", g_scaleHyperlinks =3D false, g_allowAdvOnClick =3D 1, g_showInBrowser = =3D 0, gLoopCont =3D 0, gUseSldTimings =3D 1; var g_showAnimation =3D g_supportsPPTHTML && SupportsPPTAnimation() && ( (w= indow.name=3D=3D"PPTSld" && !parent.IsFramesMode()) || g_showInBrowser );va= r g_animManager =3D null; var g_animUseRuntime =3D false; var g_animItemsToHide, g_animInteractiveItems, g_animSlideTime; var g_animMainSequence =3D null; var ENDSHOW_MESG=3D"放映结束，单击= ;鼠标退出。", SCREEN_MODE=3D"Frames", gIsEndS= how=3D0, NUM_VIS_SLDS=3D62, SCRIPT_HREF=3D"script.js", FULLSCR_HREF=3D"full= screen.htm"; var gCurSld =3D gPrevSld =3D 1, g_offset =3D 0, gNtsOpen =3D gHasNts =3D gO= tlTxtExp =3D 0, gHasNarration =3D 0, gOtlOpen =3D true window.gPPTHTML=3DSupportsPPTHTML() var gMainDoc=3Dnew Array(new hrefList("slide0002.htm",1,-1,1),new hrefList(= "slide0001.htm",1,-1,1),new hrefList("slide0003.htm",1,-1,1),new hrefList("= slide0009.htm",1,-1,1),new hrefList("slide0060.htm",1,-1,1),new hrefList("s= lide0049.htm",1,-1,1),new hrefList("slide0084.htm",1,-1,1),new hrefList("sl= ide0051.htm",1,-1,1),new hrefList("slide0016.htm",1,-1,1),new hrefList("sli= de0017.htm",1,-1,1),new hrefList("slide0061.htm",1,-1,1),new hrefList("slid= e0062.htm",1,-1,1),new hrefList("slide0018.htm",1,-1,1),new hrefList("slide= 0085.htm",1,-1,1),new hrefList("slide0019.htm",1,-1,1),new hrefList("slide0= 020.htm",1,-1,1),new hrefList("slide0063.htm",1,-1,1),new hrefList("slide00= 64.htm",1,-1,1),new hrefList("slide0065.htm",1,-1,1),new hrefList("slide001= 2.htm",1,-1,1),new hrefList("slide0066.htm",1,-1,1),new hrefList("slide0050= .htm",1,-1,1),new hrefList("slide0014.htm",1,-1,1),new hrefList("slide0015.= htm",1,-1,1),new hrefList("slide0067.htm",1,-1,1),new hrefList("slide0068.h= tm",1,-1,1),new hrefList("slide0023.htm",1,-1,1),new hrefList("slide0024.ht= m",1,-1,1),new hrefList("slide0069.htm",1,-1,1),new hrefList("slide0025.htm= ",1,-1,1),new hrefList("slide0072.htm",1,-1,1),new hrefList("slide0070.htm"= ,1,-1,1),new hrefList("slide0083.htm",1,-1,1),new hrefList("slide0026.htm",= 1,-1,1),new hrefList("slide0074.htm",1,-1,1),new hrefList("slide0073.htm",1= ,-1,1),new hrefList("slide0029.htm",1,-1,1),new hrefList("slide0028.htm",1,= -1,1),new hrefList("slide0071.htm",1,-1,1),new hrefList("slide0057.htm",1,-= 1,1),new hrefList("slide0052.htm",1,-1,1),new hrefList("slide0076.htm",1,-1= ,1),new hrefList("slide0053.htm",1,-1,1),new hrefList("slide0054.htm",1,-1,= 1),new hrefList("slide0058.htm",1,-1,1),new hrefList("slide0059.htm",1,-1,1= ),new hrefList("slide0037.htm",1,-1,1),new hrefList("slide0044.htm",1,-1,1)= ,new hrefList("slide0038.htm",1,-1,1),new hrefList("slide0039.htm",1,-1,1),= new hrefList("slide0040.htm",1,-1,1),new hrefList("slide0077.htm",1,-1,1),n= ew hrefList("slide0078.htm",1,-1,1),new hrefList("slide0079.htm",1,-1,1),ne= w hrefList("slide0045.htm",1,-1,1),new hrefList("slide0046.htm",1,-1,1),new= hrefList("slide0080.htm",1,-1,1),new hrefList("slide0048.htm",1,-1,1),new = hrefList("slide0032.htm",1,-1,1),new hrefList("slide0033.htm",1,-1,1),new h= refList("slide0081.htm",1,-1,1),new hrefList("slide0034.htm",1,-1,1)); /********************************************* Frameset functions These functions control slide navigation and state of the frameset. **********************************************/ function FullScrInit() { g_allowAdvOnClick =3D GetCurSld().mAdvOnClk document.body.style.backgroundColor=3D"black" document.oncontextmenu=3Dparent._CM; document.onkeydown =3D _KDH; document.ondragstart=3DCancel document.onselectstart=3DCancel self.focus() } function Redirect( frmId ) {=09 var str=3Ddocument.location.hash,idx=3Dstr.indexOf('#'), sId=3DGetSldId() if(idx>=3D0) str=3Dstr.substr(1); if( window.name !=3D frmId && ( sId !=3D str) ) { obj =3D GetObj("Main-File") window.location.href=3Dobj.href+"#"+sId return 1 } return 0 } var MHTMLPrefix =3D CalculateMHTMLPrefix();=20 function CalculateMHTMLPrefix() { if ( document.location.protocol =3D=3D 'mhtml:') {=20 href=3Dnew String(document.location.href)=20 Start=3Dhref.indexOf('!')+1=20 End=3Dhref.lastIndexOf('/')+1=20 if (End < Start)=20 return href.substring(0, Start)=20 else=20 return href.substring(0, End)=20 } return ''; } function GetTags(base,tag) { if(g_supportsPPTHTML) return base.all.tags(tag); else return base.getElementsByTagName(tag); } function UpdNtsPane(){ if(frames["PPTNts"]) PPTNts.location.replace( MHTMLP= refix+GetHrefObj( gCurSld ).mNtsHref ) } function UpdNavPane( sldIndex ){ if(gNavLoaded) PPTNav.UpdNav() } function UpdOtNavPane(){ if(gOtlNavLoaded) PPTOtlNav.UpdOtlNav() } function UpdOtlPane(){ if(gOtlLoaded) PPTOtl.UpdOtl() } function SetHasNts( fVal ) { if( gHasNts !=3D fVal ) { gHasNts=3DfVal UpdNavPane() } } function ToggleOtlText() { gOtlTxtExp=3D!gOtlTxtExp UpdOtlPane() } function ClearMedia() { // Clear any sounds playing before launching another browser window. Other= wise, // in fullscreen mode, you'll continue to hear the sound in the frames mod= e. if (PPTSld.pptSound) PPTSld.pptSound.loop =3D 0; } function FullScreen() {=20 if ( PPTSld.g_animUseRuntime ) PPTSld.document.body.pause(); ClearMedia(); var href =3D ( document.location.protocol =3D=3D 'mhtml:') ? FULLSCR_HREF = : FULLSCR_HREF+"#"+GetHrefObj(gCurSld).mSldHref; if(PPTNav.event.ctrlKey) { var w =3D (window.screen.availWidth * 1.0) / 2.0 var h =3D w * (PPTSld.g_origH * 1.0) / PPTSld.g_origW win =3D window.open( MHTMLPrefix+href,null,"toolbar=3D0,resizable=3D1,top= =3D0,left=3D0," + "width=3D"+ w + ",height=3D" + h ); if( win.document.body && PPTSld.g_animUseRuntime ) win.document.body.PPTSldFrameset=3Dwindow; } else { win =3D window.open( MHTMLPrefix+href,null,"fullscreen=3Dyes" ); if( win.document.body && PPTSld.g_animUseRuntime ) win.document.body.PPTSldFrameset=3Dwindow; } } function ToggleVNarration() { rObj=3DPPTSld.document.all("NSPlay") if( rObj && !PPTSld.g_animUseRuntime ) { if( (rObj.playState =3D=3D 1)||(rObj.playState =3D=3D 0) ) rObj.Play() else if( rObj.playState =3D=3D 2 ) rObj.Pause() else return; } else if( PPTSld.g_animUseRuntime ) { narObj =3D PPTSld.document.all("narrationID") if( narObj ) narObj.togglePause() } } function GetCurSldNum() { =20 obj=3DGetHrefObj(gCurSld) if( obj.mOrigVis =3D=3D 1 ) return obj.mSldIdx else =20 return gCurSld } function GetNumSlds() { =20 if( GetHrefObj(gCurSld).mOrigVis =3D=3D 1 ) return GetSldList().mNumVisSlds; else return GetSldList().mList.length } function GetSldNum( href ) { for(ii=3D0; ii 1 ) PopSldList(); else if( !IsFramesMode() ) { if( gLoopCont ) GoToFirst() else EndShow() } } function GoToPrevSld() { ii=3DgCurSld-1 if( ii > 0 ) { obj=3DGetHrefObj(ii) while ( obj && ( obj.mVis =3D=3D 0 ) && ( ii>0 ) ) obj=3DGetHrefObj(--ii) if( ii =3D=3D 0 ) ii=3D1 GoToSldNum(ii) } } function GoToFirst(){ GoToSld( GetHrefObj(1).mSldHref ) } function GoToLast() { ii=3DGetSldList().mList.length if( ii !=3D gCurSld ) GoToSld( GetHrefObj(ii).mSldHref ) } function GoToSldNum( num ) { if( PPTSld.event ) PPTSld.event.cancelBubble=3Dtrue obj =3D GetHrefObj( num ) obj.mVis=3D1 gPrevSld=3DgCurSld gCurSld =3D num; PPTSld.location.replace(MHTMLPrefix+obj.mSldHref) if( IsFramesMode() ) { UpdNavPane(); UpdOtlPane(); UpdNtsPane() } } function GoToSld( href ) { if( PPTSld.event ) PPTSld.event.cancelBubble=3Dtrue GetHrefObj( GetSldNum(href) ).mVis=3D1 PPTSld.location.replace(MHTMLPrefix+href) } function SldUpdated( id ) { if( id =3D=3D GetHrefObj(gCurSld).mSldHref ) return gPrevSld=3DgCurSld gCurSld=3DGetSldNum(id) if( IsFramesMode() ) { UpdNavPane(); UpdOtlPane(); UpdNtsPane() } } function PrevSldViewed(){ GoToSld( GetHrefObj(gPrevSld).mSldHref ) } function HasPrevSld() { return ( gIsEndShow || ( gCurSld !=3D 1 && GetHrefO= bj( gCurSld-1 ).mVis =3D=3D 1 )||( GetCurSldNum() > 1 ) ) } function HasNextSld() { return (GetCurSldNum() !=3D GetNumSlds()) } function CloseWindow() { if( HideMenu() ) return; =09 var event =3D PPTSld.event; if( !IsFramesMode() && event && (event.keyCode=3D=3D27 || event.keyCode=3D= =3D32 || event.type=3D=3D"click" ) ) window.close( self ); CatchNumKeys( self, event ); } function Unload() { gIsEndShow=3D0; } function SetupEndShow() { gIsEndShow=3D1; PPTSld.document.body.scroll=3D"no"; PPTSld.document.onkeypress=3DCloseWindow; PPTSld.document.onclick=3DCloseWindow; PPTSld.document.oncontextmenu=3D_CM; } function EndShow() { if( IsFramesMode() ) return if( PPTSld.event ) PPTSld.event.cancelBubble=3Dtrue doc=3DPPTSld.document var dir =3D doc.body.dir if( dir !=3D "rtl" ) dir =3D "ltr"; doc.open() doc.writeln('

' + ENDSHOW_MESG + '
') doc.close() } function SetSldVisited(){ GetSldList().mList[gCurSld-1].mVisited=3Dtrue } function IsSldVisited(){ return GetSldList().mList[gCurSld-1].mVisited } function hrefList( sldHref, visible, advDelay, advClk ) { this.mSldHref=3D this.mNtsHref =3D sldHref this.mOrigVis=3D this.mVis =3D visible this.mVisited=3D false this.mAdvDelay=3D advDelay this.mAdvOnClk=3D advClk } function SldList(arr,curSld,fEnd) { this.mCurSld =3D curSld; this.mList =3D new Array(); var idx =3D 1; for(ii=3D0;ii 0) { PushSldList(sldList,fEnd); gCurSld =3D 1; } else if( PPTSld.event ) PPTSld.event.cancelBubble=3Dtrue } function PushSldList(arr,fEnd) { var ii =3D gSldStack.length; gSldStack[ii] =3D new SldList(arr,gCurSld,fEnd); GoToSld( gSldStack[ii].mList[0].mSldHref ); } function PopSldList() { if (gSldStack[gSldStack.length-1].fEndShow) EndShow() else { gCurSld =3D gSldStack[gSldStack.length-1].mCurSld; gSldStack[gSldStack.length-1] =3D null; gSldStack.length--; var sldList =3D gSldStack[gSldStack.length-1]; GoToSld( sldList.mList[gCurSld - 1].mSldHref ); } } var custShowList=3Dnew Array(); /********************************************* Navigation button implementation There are 2 types of buttons: ImgBtn, TxtBtn implemented as function objects. They share a similiar interface so the event handlers can call SetActive, for example, on a button=20 object without needing to know exactly=20 what type of button it is. **********************************************/ //---------------------------------- function ImgBtn( oId,bId,w,action ) //---------------------------------- { var t=3Dthis t.Perform =3D _IBP t.SetActive =3D _IBSetA t.SetInactive=3D _IBSetI t.SetPressed =3D _IBSetP t.SetDisabled=3D _IBSetD t.Enabled =3D _IBSetE t.ChangeIcon =3D null t.UserAction =3D action t.ChgState =3D _IBUI t.mObjId =3D oId t.mBorderId=3D bId t.mWidth =3D w t.mIsOn =3D t.mCurState =3D 0 } function _IBSetA() { if( this.mIsOn ) { obj=3Dthis.ChgState( gHiliteClr,gShadowClr,2 ) obj.style.posTop=3D0 } } function _IBSetI() { if( this.mIsOn ) { obj=3Dthis.ChgState( gFaceClr,gFaceClr,1 ) obj.style.posTop=3D0=20 } } function _IBSetP() { if( this.mIsOn ) { obj=3Dthis.ChgState( gShadowClr,gHiliteClr,2 ) obj.style.posLeft+=3D1; obj.style.posTop+=3D1 } } function _IBSetD() { =20 obj=3Dthis.ChgState( gFaceClr,gFaceClr,0 ) obj.style.posTop=3D0=20 } function _IBSetE( state ) { var t=3Dthis GetObj( t.mBorderId ).style.visibility=3D"visible" if( state !=3D t.mIsOn ) { t.mIsOn=3Dstate if( state ) t.SetInactive() else t.SetDisabled() } } function _IBP() { var t=3Dthis if( t.mIsOn ) { if( t.UserAction !=3D null ) t.UserAction() if( t.ChangeIcon ) { obj=3DGetObj(t.mObjId) if( t.ChangeIcon() ) obj.style.posLeft=3Dobj.style.posLeft+(t.mCurState-4)*t.mWidth else obj.style.posLeft=3Dobj.style.posLeft+(t.mCurState-0)*t.mWidth } t.SetActive() } =20 } function _IBUI( clr1,clr2,nextState ) { var t=3Dthis SetBorder( GetObj( t.mBorderId ),clr1,clr2 ) obj=3DGetObj( t.mObjId ) obj.style.posLeft=3Dobj.style.posLeft+(t.mCurState-nextState)*t.mWidth-obj= .style.posTop t.mCurState=3DnextState return obj } //----------------------------------------- function TxtBtn( oId,oeId,action,chkState ) //----------------------------------------- { var t=3Dthis t.Perform =3D _TBP t.SetActive =3D _TBSetA t.SetInactive=3D _TBSetI t.SetPressed =3D _TBSetP t.SetDisabled=3D _TBSetD t.SetEnabled =3D _TBSetE t.GetState =3D chkState t.UserAction =3D action t.ChgState =3D _TBUI t.mObjId =3D oId t.m_elementsId=3D oeId t.mIsOn =3D 1 } function _TBSetA() { var t=3Dthis if( t.mIsOn && !t.GetState() ) t.ChgState( gHiliteClr,gShadowClr,0,0 ) } function _TBSetI() { var t=3Dthis if( t.mIsOn && !t.GetState() ) t.ChgState( gFaceClr,gFaceClr,0,0 ) } function _TBSetP() { if( this.mIsOn ) this.ChgState( gShadowClr,gHiliteClr,1,1 ) } function _TBSetD() { =20 this.ChgState( gFaceClr,gFaceClr,0,0 ) this.mIsOn =3D 0 } function _TBSetE() { var t=3Dthis if( !t.GetState() ) t.ChgState( gFaceClr,gFaceClr,0,0 ) else t.ChgState( gShadowClr,gHiliteClr,1,1 ) t.mIsOn =3D 1 } function _TBP() { var t=3Dthis if( t.mIsOn ) {=20 if( t.UserAction !=3D null ) t.UserAction() if( !t.GetState ) return if( t.GetState() ) t.SetPressed() else t.SetActive() } =20 } function _TBUI( clr1,clr2,lOffset,tOffset ) { SetBorder( GetObj( this.mObjId ),clr1,clr2 ) Offset( GetObj( this.m_elementsId ),lOffset,tOffset ) } function Offset( obj, top, left ){ obj.style.top=3Dtop; obj.style.left=3Dle= ft } function SetBorder( obj, upperLeft, lowerRight ) { s=3Dobj.style; s.borderStyle =3D "solid" s.borderWidth =3D 1=20 s.borderLeftColor =3D s.borderTopColor =3D upperLeft s.borderBottomColor=3D s.borderRightColor =3D lowerRight } function GetBtnObj(){ return gBtnArr[window.event.srcElement.id] } function BtnOnOver(){ b=3DGetBtnObj(); if( b !=3D null ) b.SetActive() } function BtnOnDown(){ b=3DGetBtnObj(); if( b !=3D null ) b.SetPressed() } function BtnOnOut(){ b=3DGetBtnObj(); if( b !=3D null ) b.SetInactive() } function BtnOnUp() { b=3DGetBtnObj() if( b !=3D null ) b.Perform() else Upd() } function GetNtsState(){ return parent.gNtsOpen } function GetOtlState(){ return parent.gOtlOpen } function GetOtlTxtState(){ return parent.gOtlTxtExp } function NtsBtnSetFlag( fVal ) { s=3Ddocument.all.item( this.m_flagId ).style s.display=3D"none" if( fVal ) s.display=3D"" else s.display=3D"none" } function _BSetA_Border(){ b =3D gBtnArr[this.mObjId]; if( b !=3D null ) b.S= etActive() } function _BSetI_Border(){ b =3D gBtnArr[this.mObjId]; if( b !=3D null ) b.S= etInactive() } function _BSetP_Border(){ b =3D gBtnArr[this.mObjId]; if( b !=3D null ) b.S= etPressed() } function _BSetA_BorderImg() {=20 b =3D gBtnArr[this.mBorderId]=20 if( b !=3D null && this.mIsOn && !b.GetState() ) { obj=3Dthis.ChgState( gHiliteClr,gShadowClr,2 ) obj.style.posTop=3D0 } } function _BSetI_BorderImg() {=20 b =3D gBtnArr[this.mBorderId] if( b !=3D null && this.mIsOn && !b.GetState() ) { obj=3Dthis.ChgState( gFaceClr,gFaceClr,1 ) obj.style.posTop=3D0 } } var gHiliteClr=3D"THREEDHIGHLIGHT",gShadowClr=3D"THREEDSHADOW",gFaceClr=3D"= THREEDFACE" var gBtnArr =3D new Array() gBtnArr["nb_otl"] =3D new TxtBtn( "nb_otl","nb_otlElem",parent.ToggleOtlPan= e,GetOtlState ) gBtnArr["nb_otlElem"] =3D new TxtBtn( "nb_otl","nb_otlElem",parent.ToggleOt= lPane,GetOtlState ) gBtnArr["nb_nts"] =3D new ImgBtn( "nb_nts","nb_ntsBorder",10,parent.ToggleN= tsPane ) gBtnArr["nb_nts"].SetActive =3D _BSetA_BorderImg; gBtnArr["nb_nts"].SetInactive =3D _BSetI_BorderImg; gBtnArr["nb_ntsBorder"] =3D new TxtBtn( "nb_ntsBorder","nb_ntsElem",parent.= ToggleNtsPane,GetNtsState ) gBtnArr["nb_ntsElem"] =3D new TxtBtn( "nb_ntsBorder","nb_ntsElem",parent.To= ggleNtsPane,GetNtsState ) gBtnArr["nb_prevBorder"] =3D gBtnArr["nb_prev"]=3D new ImgBtn( "nb_prev","n= b_prevBorder",30,parent.GoToPrevSld ) gBtnArr["nb_nextBorder"] =3D gBtnArr["nb_next"]=3D new ImgBtn( "nb_next","n= b_nextBorder",30,parent.GoToNextSld ) gBtnArr["nb_sldshw"]=3D new ImgBtn( "nb_sldshw","nb_sldshwBorder",18,parent= .FullScreen ) gBtnArr["nb_sldshwBorder"] =3D new TxtBtn( "nb_sldshw","nb_sldshwBorder",pa= rent.FullScreen,null ) gBtnArr["nb_sldshwBorder"].SetActive =3D _BSetA_Border; gBtnArr["nb_sldshwBorder"].SetInactive =3D _BSetI_Border; gBtnArr["nb_sldshwText"] =3D new TxtBtn( "nb_sldshw","nb_sldshwText",parent= .FullScreen,null ) gBtnArr["nb_sldshwText"].SetActive =3D _BSetA_Border; gBtnArr["nb_sldshwText"].SetInactive =3D _BSetI_Border; gBtnArr["nb_voice"] =3D gBtnArr["nb_voiceBorder"] =3D new ImgBtn( "nb_voice= ","nb_voiceBorder",18,parent.ToggleVNarration ) gBtnArr["nb_otlTxtBorder"] =3D gBtnArr["nb_otlTxt"]=3D new ImgBtn( "nb_otlT= xt","nb_otlTxtBorder",23,parent.ToggleOtlText ) gBtnArr["nb_ntsBorder"].m_flagId=3D "nb_nts" gBtnArr["nb_ntsBorder"].SetFlag =3D NtsBtnSetFlag gBtnArr["nb_otlTxt"].ChangeIcon=3D GetOtlTxtState /********************************************* Context menu implementation _CM() is the function that's hooked up to the oncontextmenu event. Once we're asked to show the menu, we first build it by creating DIVs on-the-fly. Then we position it=20 within the screen area so it doesn't get clipped. Creating the DIVs using createElement() means we don't have to write out any extra HTML into the slide HTML files. **********************************************/ var sNext=3D"下一步",sPrev=3D"上一步",s= End=3D"结束放映",sFont=3D"宋体",sArrow= =3D"箭头",sFreeform=3D"任意形状",sRect= =3D"矩形",sOval=3D"椭圆" function ShowMenu() { BuildMenu(); var doc=3DPPTSld.document.body,x=3DPPTSld.event.clientX+doc.scrollLeft,y= =3DPPTSld.event.clientY+doc.scrollTop m =3D PPTSld.document.all.item("ctxtmenu") m.style.pixelLeft=3Dx if( (x+m.scrollWidth > doc.clientWidth)&&(x-m.scrollWidth > 0) ) m.style.pixelLeft=3Dx-m.scrollWidth m.style.pixelTop=3Dy if( (y+m.scrollHeight > doc.clientHeight)&&(y-m.scrollHeight > 0) ) m.style.pixelTop=3Dy-m.scrollHeight m.style.display=3D"" } function _CM() { if( !parent.IsFullScrMode() ) return; if(!PPTSld.event.ctrlKey) { ShowMenu() return false } else HideMenu() } function BuildMenu() { if( PPTSld.document.all.item("ctxtmenu") ) return var mObj=3DCreateItem( PPTSld.document.body ) mObj.id=3D"ctxtmenu" mObj.style.visibility=3D"hidden" var s=3DmObj.style s.position=3D"absolute" s.cursor=3D"default" s.width=3D"120px" SetCMBorder(mObj,"menu","black") var iObj=3DCreateItem( mObj ) SetCMBorder( iObj, "threedhighlight","threedshadow" ) iObj.style.padding=3D2 CreateMenuItem( iObj,sNext,M_GoNextSld,M_True ) CreateMenuItem( iObj,sPrev,M_GoPrevSld,M_HasPrevSld ) =09 CreateSeparator( iObj ) CreateMenuItem( iObj,sEnd,M_End,M_True ) mObj.style.visibility=3D"visible" } function Cancel() { window.event.cancelBubble=3Dtrue; window.event.returnVa= lue=3Dfalse } function Highlight() { ChangeClr("activecaption","threedhighlight") } function Deselect() { ChangeClr("threedface","menutext") } function Perform() { e=3DPPTSld.event.srcElement if( e.type=3D=3D"menuitem" && e.IsActive() ) e.Action() else PPTSld.event.cancelBubble=3Dtrue } function ChangeClr( bg,clr ) { e=3DPPTSld.event.srcElement if( e.type=3D=3D"menuitem" && e.IsActive() ) { e.style.backgroundColor=3Dbg e.style.color=3Dclr } } function M_HasPrevSld() { return( parent.HasPrevSld() ) } function M_GoNextSld() { if( gIsEndShow ) M_End(); else GoToNextSld() } function M_GoPrevSld() { if( gIsEndShow ) { gIsEndShow=3D0; history.back();= PPTSld.event.cancelBubble=3Dtrue; } else GoToPrevSld() } function M_True() { return true } function M_End() { window.close( self ) } function CreateMenuItem( node,text,action,eval ) { var e=3DCreateItem( node ) e.type=3D"menuitem" e.Action=3Daction e.IsActive=3Deval e.innerHTML=3Dtext if( !e.IsActive() ) e.style.color=3D"threedshadow" e.onclick=3DPerform e.onmouseover=3DHighlight e.onmouseout=3DDeselect s=3De.style; s.fontFamily=3DsFont s.fontSize=3D"9pt" s.paddingLeft=3D2 } function CreateSeparator( node ) { var sObj=3DCreateItem( node ) SetCMBorder(sObj,"menu","menu") var s=3DsObj.style s.borderTopColor=3D"threedshadow" s.borderBottomColor=3D"threedhighlight" s.height=3D1 s.fontSize=3D"0px" } function CreateItem( node ) { var elem=3DPPTSld.document.createElement("DIV") node.insertBefore( elem ) return elem } function SetCMBorder( o,ltClr,rbClr ) { var s=3Do.style s.backgroundColor=3D"menu" s.borderStyle=3D"solid" s.borderWidth=3D1 s.borderColor=3DltClr+" "+rbClr+" "+rbClr+" "+ltClr } ------=_NextPart_01C4842D.58317ED0 Content-Location: file:///C:/85899A51/11.files/fullscreen.htm Content-Transfer-Encoding: quoted-printable Content-Type: text/html; charset="us-ascii"
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&#= 22791;注

大纲

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第十= 一章

DNA的生物合成

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五、 DNA连接酶(DNA ligase)

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复制= ;的终止

原核生物复制终止

真核生物复制终止

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一、= ;基因突变的意义

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第五= ;节   逆转录&#= 29616;象和逆转录酶

逆转ô= 05;酶（reverse transcriptase）又称为反转录= ;酶，为依赖RNA的DNA聚合%= 238;(RNA-dependent DNA polymerase, RDDP)

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= = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = = =
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2	第十= 一章 DNA的生物合成
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26	五、 DNA连接酶(DNA ligase)
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41	复制= ;的终止原核生物复制终止真核生物复制终止
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48	一、= ;基因突变的意义
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59	第五= ;节逆转录&#= 29616;象和逆转录酶逆转ô= 05;酶（reverse transcriptase）又称为反转录= ;酶，为依赖RNA的DNA聚合%= 238;(RNA-dependent DNA polymerase, RDDP)
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